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Daria Morgendorffer: The titular character of the series, Daria is present in every episode and central to almost every plot. Abrasive and cynical to a terminal degree, Daria's intellectuality and academic insight is often counterbalanced by her devout lack of motivation. Her trademark on the show was that she rarely changed the expression on her face or the tone of her voice, except for a small satisfied smile akin to the Mona Lisa's when she is pleased. Daria is voiced by Tracy Grandstaff.
Jane Lane: Daria's artistic best friend and fellow outcast, as well as the youngest of the five Lane siblings. While sharing some of Daria's cynical views and attitudes, Jane is much more lively and well adjusted than her best friend. Living with her older brother Trent, she manages to stay completely functional and content with life despite complete absence of parental guidance or even presence. Jane is the show's second central character, and is voiced by Wendy Hoopes.
Together: Daria and Jane initially meet in a self-esteem class in the very first episode of the series, but they are quick to form a very close friendship. Throughout the series, Jane and Daria spend most of their together, both in and out of school. Their friendship is the vehicle for most the progression in the storylines. Daria and Jane share a mutual pessimistic view on the world, the two are also highly artistic (Daria in her writing and Jane in her paintings/sketches). They also love pizza, and visit their local pizza restaurant regularly.